The issue may be with the 126, or it may be with your driver. Or both. If you have a volt meter, measure the voltage on the direction signal (J5 A or B, which ever it is). Jog one direction (then stop) and then measure the voltage on the PMDX-126 screw terminal. Then jog the other way (and stop) and then measure the voltage again. It should change from close to zero volts for one direction to close to 5 volts for the other.
If you don't see those voltages, disconnect the step/dir wires from the 126 and repeat the experiment. Do you now see close to 0 and 5 volts?
Try connecting one of the other motor drivers to the same 126 pins (ex. J5 A and B) and see if it works. Then try the known good driver on the original 126 pins (2 and 3).